Jambo was the first male silverback gorilla to be born in captivity, the first to be mother-reared and was one of the most prolific breeding silverbacks.

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He is charged with four counts of kidnapping, covering the three initial abduction victims and Jocelyn, Ms Berry's six-year-old daughter, who was apparently conceived and born in captivity.

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Ms Berry, 27, arrived at her sister's home shortly before midday on Wednesday, along with her six-year-old daughter Jocelyn, who is understood to have been conceived and born in captivity.

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The situation is likely also difficult for Dugard's two daughters, born to her in captivity, who have never met Dugard's family before, Napoletano said.

CNN: For long-separated families, reunions can be a struggle