However, genetic studies revealed that there are actually two quite distinct clouded leopard species.

BBC: Earth News - Clouded leopard: First film of new Asia big cat species

The scientists suspect a large male clouded leopard made the kill, and had removed part of the front right leg.

BBC: Earth News - Clouded leopard: First film of new Asia big cat species

Earlier this year, the same project released the first video to be made public of a wild Sundaland clouded leopard.

BBC: Earth News - Rare Otter civet filmed for first time in Borneo

During the surveys, the research team also discovered a juvenile samba deer (Cervus unicolor) which had been killed by a clouded leopard.

BBC: Earth News - Clouded leopard: First film of new Asia big cat species

This leopard is now known as the Sunda or Sundaland clouded leopard (Neofelis diardi), though it was previously and erroneously called the Bornean clouded leopard.

BBC: Earth News - Clouded leopard: First film of new Asia big cat species

And a tourist is thought to have taken a 30 second video of a wild Sundaland clouded leopard in 2006, but that video has never been made public.

BBC: Earth News - Clouded leopard: First film of new Asia big cat species

The clouded leopard, the largest predator on Borneo, appears to live at very low densities within the reserve, as it has only rarely been photographed by the researchers or camera traps.

BBC: Earth News - Clouded leopard: First film of new Asia big cat species

As well as the better known clouded leopard living on the Asian mainland (Neofelis nebulosa), scientists determined that a separate clouded leopard species lives on the islands of Borneo and Sumatra.

BBC: Earth News - Clouded leopard: First film of new Asia big cat species