Investigating the evolution of the lion social structure in such detail, Prof Packer and his team have had to look closely at the many theories for this lifestyle that have been suggested over the years, such as co-operative hunting, communal suckling of cubs and defence of young against attack.

BBC: Lions breed better if living near joining rivers

While they may seem attractive theories on the surface, on closer inspection the research team found they could not be the driving force in the evolution of the lion's social lifestyle.

BBC: Lions breed better if living near joining rivers

They deepen their involvement with the Suns through service opportunities, cultural activities like traditional Chinese lion dances and frequent social events that take them out of their comfort zones and expand their circles.

WHITEHOUSE: Once a Sun, Always a Sun

He was the lion of the international movement in social-realist public art, which peaked in the nineteen-thirties.

NEWYORKER: The Painting on the Wall