And, with Stewart Downing's crossing completely ineffective on the opposite wing, England quickly ran out of ideas.
Winger Stewart Downing was almost lost for words after playing his part in the amazing comeback.
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And moments later, Jaaskelainen was again alert to deny Stewart Downing's drive from 20 yards out.
Steven Gerrard went close, Pepe Reina denied Stewart Downing and Gabriel Agbonlahor, and John Carew headed wide.
Villa have also added Stewart Downing, Fabian Delph, Habib Beye and Andy Marshall to their squad this summer.
He made eight changes to the team, with Ashley Young, Stewart Downing and Marc Albrighton named as substitutes.
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Stewart Downing gave the Reds the lead in the second minute before the Uruguayan striker took center stage.
They went ahead when Stewart Downing sliced Gary McSheffrey's free-kick into his own net inside the six-yard box.
Stewart Downing was the rare exception and provided the few moments of class.
Capello has called up Stewart Downing for the first time in 11 months but Chelsea midfielder Joe Cole has been overlooked.
Villa boss Martin O'Neill was pondering the introduction of Stewart Downing when the outstanding Malouda made it five after 68 minutes.
Stewart Downing's cross caused confusion, and Vidic was lucky to see Heskey's header bounce off his shoulder on to the bar.
Phil Bardsley was yellow-carded for almost rugby-tackling Stewart Downing and from the resultant free-kick Rochemback unleashed a fierce drive from 25 yards.
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Bent then headed home from Stewart Downing's fine right-wing cross but the goal was harshly disallowed for a push on Lars Jacobsen.
It was thought that Premier League vultures would swoop for the club's assets such as David Wheater, Tuncay and the injured Stewart Downing.
Middlesbrough threatened from the flanks, with Stewart Downing and Tuncay both sending in dangerous crosses that just failed to result in openings for Alves.
England midfielder Stewart Downing, Egyptian striker Mido and goalkeeper Ross Turnbull were the only high-profile names to leave the Riverside during the close season.
Boro's academy has unearthed several notable homegrown players in recent years, including England international Stewart Downing, and under-21 stars Andrew Taylor, Lee Cattermole and David Wheater.
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The dramatic turn in fortunes forced McClaren into a triple substitution, sending on Peter Crouch, Frank Lampard and Stewart Downing for Lescott, Cole and Shaun Wright-Phillips.
They threatened to open the scoring inside a minute when Marvin Emnes released Stewart Downing and his cross had to be poked behind by Gareth Barry.
The club have already lost Stewart Downing, who joined Aston Villa last week, and strikers Mido and Tuncay are also thought to be heading for the exit.
The brave but tiring visitors had no riposte and Villa continued to pile forward, with Speroni performing more heroics to superbly tip over Stewart Downing's 20-yard drive.
Redknapp is also keen to sign a left-sided midfielder in the January transfer window but Middlesbrough have already rejected two bids from the Londoners for Stewart Downing.
Stewart Downing broke well and combined with Sanli Tuncay before feeding O'Neil, who took a touch and tucked the ball in at the near post with aplomb.
He was replaced by Stewart Downing, while Ben Foster replaced James and Beckham was able to add another cap to his collection at the expense of Lennon.
It was Villa who created the best late chances and only two fine saves by Robinson stopped substitute Stewart Downing from adding to their tally before the end.
Wheater headed wide at the other end when Duke failed to punch a Stewart Downing free-kick away before an escape for the home side at the other end.
As Boro regrouped United were indebted to some brave defensive play from Wes Brown, who first tackled Yakubu and then came across to block Stewart Downing's attempted cross.
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Stiliyan Petrov then made a bizarre decision to cross the ball, almost away from goal, when presented with a shooting chance in the box following Stewart Downing's intelligent pass.
The Eagles were on the verge of an unlikely fifth-round victory when Petrov plunged to meet Stewart Downing's corner with a diving header to hand Martin O'Neill's side an 87th-minute equaliser.
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