An adult male lion is resting in the grass on the Kenyan savanna.

肯尼亚热带草原上，一头成年雄狮正在草丛里休息。

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A decade later another study reported that adult male white-tailed deer exhibited forehead rubbing just before and during the rut.

十年后的另一项研究报告称，成年雄性白尾鹿在即将发情和发情期间会摩擦前额。

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Adult male deer usually produce rubs in late summer or early autumn when the outer velvet layer is being shed from their antlers.

成年公鹿通常在夏末或初秋鹿茸脱落时制造擦痕。

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Adult male deer usually produce rubs in late summer or early autumn when the outer velvet layer is being shed from their antlers.

成年公鹿通常在夏末或初秋鹿茸脱落时制造擦痕。

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