However only indirect approaches can be applied to the determination of oxygen demanded by tiny living individuals because in this case it is hard to make direct measurement.
然而对于一些微小生物个体无法直接测量，只有通过间接的方法求得。
However only indirect approaches can be applied to the determination of oxygen demanded by tiny living individuals because in this case it is hard to make direct measurement.
然而对于一些微小生物个体无法直接测量，只有通过间接的方法求得。
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