He gave the name of the chief culprit.

他供出了主犯的名字。

《新英汉大辞典》

The chief culprit is Germany.

罪魁祸首是德国。

youdao

And triglycerides high is the chief culprit of vascular sclerosis.

而甘油三脂水平高正是血管硬化的罪魁祸首。

youdao

The chief culprit appears to be his declining LD%, with corresponding increases in his GB% and FB%.

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It is true that nuclear energy does not produce carbon dioxide, the chief culprit behind man-made global warming.

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That would fix what he seems to think was the chief culprit in the crisis: how badly everyone underestimated risk.

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