Dr Federle and his colleagues argue that the complex arrangement of pegs and channels around the hexagonal cells has evolved to drain the mucus rapidly away from the toepads, allowing the tree frog to grip.
They calculated the force in question from the contact area of the toe, and found that toepads were still sticky two minutes after the sliding had stopped, long after the point when, according to their estimate of its viscosity, all the mucus would have drained away.