The Scottish Daily Express sells around 60, 000 copies a day - more than either The Herald or The Scotsman but only around half the sale of its direct rival the Scottish Daily Mail.

BBC: More jobs are cut at Scottish newspapers

The best performing daily paper in the Scottish market was the Daily Mail, with its December 2011 sales higher than the year before, at 107, 400.

BBC: Scottish newspaper sales plunge

Scottish newspapers have also faced an additional challenge as some UK-wide titles, especially the Daily Mail, made significant inroads into the Scottish market for the first time.

BBC: Printing press

Proposed cuts to the number of journalists at the Daily Record and Sunday Mail newspapers were raised in the Scottish Parliament yesterday, at First Minister's Questions.

BBC: What now for Scotland's newspapers?