On Saturday, a team of zoo keepers and the zoo vet were able to retrieve the escapee from a tree within the zoo grounds.

BBC: Missing coati returned to Belfast Zoo

The procedure was carried out in Vietnam by Edinburgh Zoo vet Romain Pizzi, the only UK vet specialising in zoo and wildlife keyhole surgery.

BBC: Pioneering surgery for moon bear by Edinburgh vet

Lisa Argilla, a vet at Wellington Zoo, said they suspected it had suffered multiple organ failure.

BBC: Lost Antarctic penguin dies in New Zealand zoo

Lisa Argilla, a vet at Wellington Zoo, said the penguin had possibly struggled to find enough food or had had problems hunting and had come ashore as he needed to go through his seasonal moulting.

BBC: Lost Antarctic Royal penguin found in New Zealand