Other reasons included a severe climate, and loss of living space and water to farm animals.
VOA: special.2010.06.22
High levels of domoic acid in the sardines may have worsened the effects of oxygen loss in the water as the fish crowded into a limited space.
VOA: special.2011.03.15
They also help prevent soil loss and protect water supplies.
VOA: special.2009.09.01
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